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Seventeen-year-old Mary Katherine, or M.K., (Amanda Seyfried) moves in with her eccentric scientist father, Professor Bomba (Jason Sudeikis), who has been searching for tiny human soldiers called Leafmen. They protect the forest Bomba lives near from evil creatures called Boggans and their malevolent leader Mandrake (Christoph Waltz). An independent young soldier, Nod (Josh Hutcherson) decides to quit, much to the ire of the no-nonsense Leafmen leader Ronin (Colin Farrell).
The queen of the forest, Queen Tara (Beyoncé Knowles), must choose an heir to her throne and goes out to a field of leaf pods, guarded by a laid-back slug named Mub (Aziz Ansari) and a wannabe Leafman snail named Grub (Chris O'Dowd). Tara chooses the smallest pod as the heir. Immediately after she does so, the Boggans attack. Tara flees the area with the pod, and though her bodyguards do their best to protect her, they are soon overwhelmed by the sheer number of Boggans. Tara uses her magic to slow down her pursuers, and is at one point rescued by a flower child who idolizes her. Eventually, Ronin arrives for her and the pair fly off on Ronin's hummingbird mount. They are then attacked by Mandrake and his son Dagda (Blake Anderson). Dagda is killed by Ronin, but Tara is fatally wounded when Mandrake shoots her with an arrow.
Songwriter Roger Radcliffe lives in a bachelor flat in London, England along with his dalmatian Pongo. Bored with bachelor life, Pongo decides to find a wife for Roger and a mate for himself. While watching various female dog-human pairs out the window, he spots the perfect couple, a woman named Anita and her female dalmatian, Perdita. He quickly gets Roger out of the house and drags him through the park to arrange a meeting. Pongo accidentally causes both Roger and Anita to fall into a pond, but it works out well as the couple falls in love. Both couples marry.
Later, Perdita gives birth to 15 puppies. One almost dies, but Roger is able to revive it by rubbing it in a towel. That same night, they are visited by Cruella De Vil, a wealthy and materialistic former schoolmate of Anita's. She offers to buy the entire litter for a large sum, but Roger says they are not for sale. Weeks later, she hires Jasper and Horace Badun to steal them. When Scotland Yard is unable to determine the thieves or find the puppies, Pongo and Perdita use the "Twilight Bark", normally a canine gossip line, to ask for help from the other dogs in London.
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Iranian activists storm the United States embassy in Tehran on November 4, 1979, in retaliation for President Jimmy Carter giving the Shah asylum in the U.S. during the Iranian Revolution. More than 50 of the embassy staff are taken as hostages, but six escape and hide in the home of the Canadian ambassador Ken Taylor. With the escapees' situation kept secret, the U.S. State Department begins to explore options for exfiltrating them from Iran. Tony Mendez, a U.S. Central Intelligence Agency exfiltration specialist is brought in for consultation. He criticizes the proposals, but is at a loss when it comes time to propose a rescue, as an alternative. While on the phone with his son, he is inspired by watching Battle for the Planet of the Apes and begins plans for creating a cover story for the escapees: that they are Canadian filmmakers who happened to be in Iran scouting exotic locations for a similar science-fiction film.

Mendez and his supervisor Jack O'Donnell contact John Chambers,