Seventeen-year-old Mary Katherine, or M.K., (Amanda Seyfried) moves in with her eccentric scientist father, Professor Bomba (Jason Sudeikis),
who has been searching for tiny human soldiers called Leafmen. They
protect the forest Bomba lives near from evil creatures called Boggans
and their malevolent leader Mandrake (Christoph Waltz). An independent young soldier, Nod (Josh Hutcherson) decides to quit, much to the ire of the no-nonsense Leafmen leader Ronin (Colin Farrell).
The queen of the forest, Queen Tara (Beyoncé Knowles), must choose an heir to her throne and goes out to a field of leaf pods, guarded by a laid-back slug named Mub (Aziz Ansari) and a wannabe Leafman snail named Grub (Chris O'Dowd). Tara chooses the smallest pod as the heir. Immediately after she does so, the Boggans attack. Tara flees the area with the pod, and though her bodyguards do their best to protect her, they are soon overwhelmed by the sheer number of Boggans. Tara uses her magic to slow down her pursuers, and is at one point rescued by a flower child who idolizes her. Eventually, Ronin arrives for her and the pair fly off on Ronin's hummingbird mount. They are then attacked by Mandrake and his son Dagda (Blake Anderson). Dagda is killed by Ronin, but Tara is fatally wounded when Mandrake shoots her with an arrow.
The queen of the forest, Queen Tara (Beyoncé Knowles), must choose an heir to her throne and goes out to a field of leaf pods, guarded by a laid-back slug named Mub (Aziz Ansari) and a wannabe Leafman snail named Grub (Chris O'Dowd). Tara chooses the smallest pod as the heir. Immediately after she does so, the Boggans attack. Tara flees the area with the pod, and though her bodyguards do their best to protect her, they are soon overwhelmed by the sheer number of Boggans. Tara uses her magic to slow down her pursuers, and is at one point rescued by a flower child who idolizes her. Eventually, Ronin arrives for her and the pair fly off on Ronin's hummingbird mount. They are then attacked by Mandrake and his son Dagda (Blake Anderson). Dagda is killed by Ronin, but Tara is fatally wounded when Mandrake shoots her with an arrow.